This is actually fascinating because it demonstrates a social construct created by animals. Not all animals have the same intelligence, and in nature intelligence is typically found among animals living in groups. It requires intelligence to live and cooperate with others. To me it seems that these animals actually used their intelligence to preserve socially constructed zones for the general good of multiple groups of wolves, likely by staying where the right smell is and not going where the wrong smell is. Hopefully none of the wolves discover the prisoner's dilemma and become a wolf emperor by suddenly invading and enslaving their unprepared neighbors.